Atherton, chairman of Ospreys Rugby and its holding company Llandarcy Park (Ospreys) Ltd, will be replaced by Roger Blyth, who steps up from his previous position as managing director of both companies.
Andrew Hore, the chief operations officer, will become chief executive of the two companies.
An Ospreys statement said Hore's new role would give the New Zealander "total responsibility for all operations within the Ospreys, including rugby, finance, commercial, sales and marketing, and all the other functions which make up the complete business."
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